I'm using my own Theme, i'v read this references. For comments I'm using comments.php template which is located in my theme folder, but there isn't code in comments.php which create navigation links, actually between header and main content.
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First, it would be very helpful to know what Theme you're using, so that we can give specific instructions/advice. Without knowing what Theme you're using, we have no way of knowing what template file(s) to tell you to edit; however, those links are likely coming from these two tags:
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